Texas Beetle Resources


Lady Beetles ("Ladybugs") of Texas

Family Coccinellidae

156 species of coccinellids have been recorded from Texas
Over 50% of Texas' species are illustrated here

Speciemens primarily shot at the TAMUIC and UTIC
Thumbnails link to larger photo on BugGuide.Net. Click on "Info" tab for more information.


Genus Cephaloscymnus

Cephaloscymnus sp. - Cephaloscymnus occidentalis
     Cephaloscymnus occidentalis Horn 
     Cephaloscymnus zimmermanni australis Gordon
     Cephaloscymnus sp.

Genus Delphastus  

Delphastus pusilla (LeConte) - Delphastus pusillus Delphastus pusilla (LeConte)  - Delphastus pusillus Delphastus pusilla (LeConte)  - Delphastus pusillus
     Delphastus pusillus (LeConte) 

Genus Gnathoweisea

Gnathoweisea sp., prob. texana Gordon - Gnathoweisea texana
     Gnathoweisea texana Gordon       

Genus Microweisea

Microweisea sp. - Microweisea
     Microweisea sp.
     Microweisea minuta (Casey) 

Genus Brachiacantha 

Brachiacantha barberi Gordon - Brachiacantha barberi Brachiacantha barberi Gordon - Brachiacantha barberi Brachiacantha barberi Gordon - Brachiacantha barberi
    Brachiacantha barberi
Gordon 

Brachiacantha bollii Crotch - Brachiacantha bollii - female Brachiacantha bollii Crotch - Brachiacantha bollii - female Brachiacantha bollii Crotch - Brachiacantha bollii - female Brachiacantha bollii Crotch - Brachiacantha bollii Brachiacantha bollii Crotch - Brachiacantha bollii 
     Brachiacantha bollii Crotch 

Brachiacantha decora Casey - Brachiacantha decora
     Brachiacantha decora Casey 

Brachiacantha quadrillum LeConte - Brachiacantha quadrillum Brachiacantha quadrillum LeConte - Brachiacantha quadrillum Brachiacantha quadrillum LeConte - Brachiacantha quadrillum
Brachiacantha quadrillum LeConte - Brachiacantha quadrillum Brachiacantha quadrillum LeConte - Brachiacantha quadrillum Brachiacantha quadrillum LeConte - Brachiacantha quadrillum Brachiacantha quadrillum LeConte - Brachiacantha quadrillum
     Brachiacantha quadrillum LeConte 

Brachiacantha subfasciata Mulsant - Brachiacantha subfasciata - female
     Brachiacantha subfasciata Mulsant 

Back and orange
 beetle - Brachiacantha tau Back and orange
 beetle - Brachiacantha tau
     Brachiacantha tau LeConte 

Brachiacantha testudo Casey - Brachiacantha testudo Brachiacantha testudo Casey - Brachiacantha testudo
     Brachiacantha testudo Casey 

Genus Cryptolaemus

          Mealybug Destoryer - Cryptolaemus montrouzieri - male
                Cryptolaemus montrouzieri Mulsant

Genus Didion

Histerid? - Didion punctatum Unknown  - Didion punctatum
    Didion punctatum (Melsheimer)

Genus Diomus 

Unknown Beetle - Diomus amabilis Unknown Beetle - Diomus amabilis
     Diomus amabilis (LeConte) 

Diomus debilis (LeConte) - Diomus debilis Diomus debilis (LeConte) - Diomus debilis Diomus debilis (LeConte) - Diomus debilis
     Diomus debilis (LeConte) 
     Diomus liebecki (Horn) 

Diomus pseudotaedatus Gordon - Diomus pseudotaedatus Diomus pseudotaedatus Gordon - Diomus pseudotaedatus
     Diomus pseudotaedatus Gordon 

Diomus terminatus (Say) - Diomus terminatus Diomus terminatus (Say) - Diomus terminatus Diomus terminatus (Say) - Diomus terminatus
     Diomus terminatus (Say) 
     Diomus texanus Gordon

Diomus xanthaspis (Mulsant) - Diomus xanthaspis
     Diomus xanthaspis (Mulsant) 

Genus Nephaspis 

Dusky Lady Beetle - Nephaspis oculatus Nephaspis oculatus (Blatchley) - Nephaspis oculatus Nephaspis oculatus (Blatchley) - Nephaspis oculatus
     Nephaspis oculatus (Blatchley) 

Genus Nephus 

Nephus flavifrons (Melsheimer) - Nephus flavifrons Nephus flavifrons (Melsheimer) - Nephus flavifrons Nephus flavifrons (Melsheimer) - Nephus flavifrons Nephus flavifrons (Melsheimer) - Nephus flavifrons
     Nephus flavifrons (Melsheimer) 
     Nephus guttulatus (LeConte) 

Nephus intrusus (Horn) - Nephus intrusus
     Nephus intrusus (Horn) 
     Nephus sordidus (Horn) 

Genus Helesius 

Helesius nubilans Casey - Helesius nubilans Helesius nubilans Casey - Helesius nubilans
     Helesius nubilans Casey 

Genus Hyperaspis 

Hyperaspis - female Hyperaspis - female Hyperaspis - female
     Hyperaspis sp.

Hyperaspis bensonica Casey - Hyperaspis bensonica Hyperaspis bensonica Casey - Hyperaspis bensonica Hyperaspis bensonica Casey - Hyperaspis bensonica
     Hyperaspis bensonica Casey 

 
     Hyperaspis bigeminata (Randall) 

Hyperaspis Hyperaspis
     Hyperaspis caseyi Gordon 

Ladybird Beetle - Hyperaspis connectens - female Hyperaspis connectens (Thunberg, 1808) - Hyperaspis connectens - female Ladybird Beetle - Hyperaspis connectens
     Hyperaspis connectens (Thunberg) 

hyper - Hyperaspis conspirans hyper - Hyperaspis conspirans hyper - Hyperaspis conspirans
     Hyperaspis conspirans Casey 

Hyperaspis cruenta LeConte - Hyperaspis cruenta Hyperaspis cruenta LeConte - Hyperaspis cruenta Hyperaspis cruenta LeConte - Hyperaspis cruenta
     Hyperaspis cruenta LeConte 

Hyperaspis gemma Casey - Hyperaspis gemma Hyperaspis gemma Casey - Hyperaspis gemma Hyperaspis gemma Casey - Hyperaspis gemma Hyperaspis gemma Casey - Hyperaspis gemma
     Hyperaspis gemma Casey 

Hyperaspis globula Casey - Hyperaspis globula Hyperaspis globula Casey - Hyperaspis globula Hyperaspis globula Casey - Hyperaspis globula Hyperaspis globula Casey - Hyperaspis globula
     Hyperaspis globula Casey 

Hyperaspis imitator Gordon - Hyperaspis imitator Hyperaspis imitator Gordon - Hyperaspis imitator Hyperaspis imitator Gordon - Hyperaspis imitator Hyperaspis imitator Gordon - Hyperaspis imitator
     Hyperaspis imitator Gordon 

          Hyperaspis inedita Mulsant - Hyperaspis inedita Hyperaspis inedita Mulsant - Hyperaspis inedita
               Hyperaspis inedita Mulsant - recorded from LA & OK

Hyperaspis inflexa Casey - Hyperaspis inflexa Hyperaspis inflexa Casey - Hyperaspis inflexa Hyperaspis inflexa Casey - Hyperaspis inflexa Hyperaspis inflexa Casey - Hyperaspis inflexa Hyperaspis lateralis Mulsant  - Hyperaspis inflexa Hyperaspis lateralis Mulsant  - Hyperaspis inflexa
     Hyperaspis inflexa Casey 

 Coccinellidae  - Hyperaspis lateralis - male Hyperaspis lateralis Mulsant  - Hyperaspis lateralis - female Hyperaspis lateralis Mulsant  - Hyperaspis lateralis - female Hyperaspis lateralis Mulsant  - Hyperaspis lateralis - female Hyperaspis lateralis Mulsant  - Hyperaspis lateralis - female
     Hyperaspis lateralis Mulsant 

Hyperaspis - Hyperaspis levrati Hyperaspis - Hyperaspis levrati
     Hyperaspis levrati
(Mulsant) 

Hyperaspis lugubris (Randall) - Hyperaspis lugubris Hyperaspis lugubris (Randall) - Hyperaspis lugubris
     Hyperaspis lugubris (Randall) - recorded from southeasternmost county in NM

Hyperaspis medialis Casey - Hyperaspis medialis Hyperaspis medialis Casey - Hyperaspis medialis Hyperaspis medialis Casey - Hyperaspis medialis
     Hyperaspis medialis Casey 

Hyperaspis octonotata Casey - Hyperaspis octonotata Hyperaspis octonotata Casey - Hyperaspis octonotata Hyperaspis octonotata Casey - Hyperaspis octonotata Hyperaspis octonotata Casey - Hyperaspis octonotata
     Hyperaspis octonotata Casey 

Hyperaspis pleuralis Casey - Hyperaspis pleuralis Hyperaspis pleuralis Casey - Hyperaspis pleuralis
    Hyperaspis pleuralis Casey 
    Hyperaspis pratensis LeConte

Hyperaspis proba (Say) - Hyperaspis proba Lady Beetle - Hyperaspis proba Lady beetle - Hyperaspis proba - female
    Hyperaspis proba (Say)

Hyperaspis punctata LeConte - Hyperaspis punctata Hyperaspis punctata LeConte - Hyperaspis punctata Hyperaspis punctata LeConte - Hyperaspis punctata Hyperaspis punctata LeConte - Hyperaspis punctata
     Hyperaspis punctata
LeConte 

Hyperaspis rotunda Casey - Hyperaspis rotunda Hyperaspis rotunda Casey - Hyperaspis rotunda Hyperaspis rotunda Casey - Hyperaspis rotunda Hyperaspis rotunda Casey - Hyperaspis rotunda
     Hyperaspis rotunda Casey 

Hyperaspis schaefferi Gordon - Hyperaspis schaefferi Hyperaspis schaefferi Gordon - Hyperaspis schaefferi
     Hyperaspis schaefferi Gordon 

Hyperaspis signata bicentralis Casey - Hyperaspis signata Hyperaspis signata bicentralis Casey - Hyperaspis signata Hyperaspis signata bicentralis Casey - Hyperaspis signata Hyperaspis signata bicentralis Casey - Hyperaspis signata
     Hyperaspis signata (Olivier) 
     Hyperaspis triangulum Casey 

Hyperaspis trifurcata Schaeffer - Hyperaspis trifurcata Hyperaspis trifurcata Schaeffer - Hyperaspis trifurcata Hyperaspis trifurcata Schaeffer - Hyperaspis trifurcata Hyperaspis trifurcata Schaeffer - Hyperaspis trifurcata
     Hyperaspis trifurcata Schaeffer 

Hyperaspis weisei Schaeffer - Hyperaspis weisei Hyperaspis weisei Schaeffer - Hyperaspis weisei Hyperaspis weisei Schaeffer - Hyperaspis weisei
      Hyperaspis weisei Schaeffer 

Hyperaspis wickhami Casey - Hyperaspis wickhami Hyperaspis wickhami Casey - Hyperaspis wickhami Hyperaspis wickhami Casey - Hyperaspis wickhami Hyperaspis wickhami Casey - Hyperaspis wickhami
     Hyperaspis wickhami Casey 

Genus Hyperaspidius

Hyperaspidius
    Hyperaspidius sp.

Hyperaspidius nanellus Gordon - Hyperaspidius nanellus
    Hyperaspidius nanellus Gordon
    Hyperaspidius marginatus (Gaines)

Hyperaspidius oblongus Casey - Hyperaspidius oblongus
    Hyperaspidius oblongus Casey

Genus Selvadius 

Selvadius sp. - Selvadius
     Selvadius sp.
     Selvadius rectus Casey 

Genus Scymnus 

Scymnus sp. - Scymnus
     Scymnus sp.
     Scymnus (Pullus) ardelio Horn
     Scymnus (Pullus) aridus Casey 


     Scymnus (Pullus) brullei
Mulsant 


     Scymnus (Pullus) caudalis LeConte

Scymnus cervicalis Mulsant  - Scymnus cervicalis Scymnus cervicalis Mulsant - Scymnus cervicalis
     Scymnus (Pullus) cervicalis Mulsant  
     Scymnus (Pullus) cockerelli Casey 


     Scymnus (Pullus) compar Casey

Scymnus creperus Mulsant - Scymnus creperus Scymnus creperus Mulsant - Scymnus creperus
     Scymnus (Pullus) creperus Mulsant

Scymnus enochrus Gordon - Scymnus enochrus
     Scymnus (Pullus) enochrus Gordon 
     Scymnus (Pullus) horni Gorham 
     Scymnus (Pullus) howdeni Gordon 

 Scymnus kansanus? - Scymnus kansanus  Scymnus kansanus? - Scymnus kansanus
     Scymnus (Pullus) kansanus Casey 

Scymnus (Pullus) loewii Mulsant - Scymnus loewii Scymnus loewii Mulsant - Scymnus loewii
     Scymnus (Pullus) loewii Mulsant

Scymnus louisianae J. Chapin - Scymnus louisianae Scymnus louisianae J. Chapin - Scymnus louisianae
     Scymnus (Pullus) louisianae J. Chapin

Scymnus pacificus Crotch - Scymnus pacificus Scymnus pacificus Crotch - Scymnus pacificus
     Scymnus (Pullus) pacificus Crotch 
     Scymnus (Pullus) pallens LeConte 
     Scymnus (Pullus) pauculus Gordon 

flood debris #9 - Scymnus rubricaudus flood debris #9 - Scymnus rubricaudus
     Scymnus (Pullus) rubricaudus Casey 
     Scymnus (Pullus) socer LeConte 
     Scymnus (Pullus) tenebrosus Mulsant 
     Scymnus (Pullus) uncus Wingo 
     Scymnus (Scymnus) apicanus apicanus J. Chapin

Scymnus circumspectus Horn - Scymnus circumspectus Scymnus circumspectus Horn - Scymnus circumspectus
     Scymnus (Scymnus) circumspectus Horn

scymnine - Scymnus indianensis scymnine - Scymnus indianensis
     Scymnus (Scymnus) indianensis Weise
     Scymnus (Scymnus) paracanus paracanus J. Chapin 

Genus Stethorus 

Stethorus sp. - Stethorus
     Stethorus sp.
     Stethorus caseyi Gordon & Chapin 
     Stethorus pinachi Gordon & Chapin 
     Stethorus utilis (Horn) 

Genus Thalassa

Twice- stabbed ladybeetle? - Thalassa montezumae - female Thalassa montezumae Mulsant - Thalassa montezumae - male Large Ladybird Beetle - Thalassa montezumae - male
     Thalassa montezumae Mulsant 

Genus Zagloba 

Zagloba hystrix Casey - Zagloba hystrix
     Zagloba hystrix Casey 
     Zagloba satana Gordon 


Subfamily Chilocorinae

       Genus Axion 

 Axion plagiatum - Axion plagiatum Axion venter, edited for detail - Axion plagiatum Beetle larva - Axion 
plagiatum
     Axion plagiatum (Olivier)

Three-spotted Lady Beetle - Axion tripustulatum Axion tripustulatum
     Axion tripustulatum (DeGeer)

Genus Chilocorus 

Chilocorus cacti (Linnaeus) - Chilocorus cacti Chilocorus cacti (Linnaeus) - Chilocorus cacti Chilocorus cacti (Linnaeus) - Chilocorus cacti
     Chilocorus cacti (Linnaeus) 

Twice-stabbed ladybird - Chilocorus stigma Twice-stabbed lady beetle - Chilocorus stigma Chilocorus stigma (Say) - Chilocorus stigma
     Chilocorus stigma (Say)  

Genus Exochomus 

Exochomus childreni Mulsant  - Exochomus childreni - male Exochomus childreni guexi LeConte - Exochomus childreni Exochomus childreni guexi LeConte - Exochomus childreni Exochomus childreni guexi LeConte - Exochomus childreni Exochomus childreni guexi LeConte - Exochomus childreni Exochomus childreni guexi LeConte  - Exochomus childreni
     Exochomus childreni guexi LeConte 

Dark Lady Beetle with 2 orange spots and rim around edges - Exochomus subrotundus blackish Lady Beetle with reddish markings on Goosefoot - Exochomus subrotundus Dark Lady Beetle with 2 orange spots and rim around edges - Exochomus subrotundus Dark Lady Beetle with 2 orange spots and rim around edges - Exochomus subrotundus
     Exochomus subrotundus Casey


Genus Anovia

               Anovia virginalis (Wickham)

Genus Rhyzobius

Rhyzobius lophanthae (Blaisdell) - Rhyzobius lophanthae Rhyzobius lophanthae (Blaisdell) - Rhyzobius lophanthae Rhyzobius lophanthae maybe - Rhyzobius lophanthae

                Rhyzobius lophanthae (Blaisdell) - Non-native

Genus Rodolia 

Rodolia cardinalis (Mulsant) - Rodolia cardinalis Rodolia cardinalis (Mulsant) - Rodolia cardinalis Rodolia cardinalis (Mulsant) - Rodolia cardinalis 
     Rodolia cardinalis (Mulsant) - Non-native


Subfamily Coccinellinae

       Genus Paranaemia

          Coccinelidae - Paranaemia vittigera
                Paranaemia vittigera (Mannerheim)

Genus Coccinella 

  Seven-spotted Lady Beetle larva - Coccinella septempunctata
     Coccinella septempunctata Linnaeus - Non-native

Genus Coleomegilla 

Coleomegilla maculata (DeGeer) - Coleomegilla maculata BG2558 E1670 - Coleomegilla maculata BG2558 E1670 - Coleomegilla maculata Ladybeetle larva - Coleomegilla maculata
     Coleomegilla maculata (Degeer) 

Genus Cycloneda 

Cycloneda munda (Say) - Cycloneda munda Cycloneda munda ? - Cycloneda munda Cycloneda munda (Say) - Cycloneda munda larva - Cycloneda munda
     Cycloneda munda (Say) 

Cycloneda sanguinea (Linnaeus) - Cycloneda sanguinea Cycloneda sanguinea (Linnaeus) - Cycloneda sanguinea Coccinellid Larva -
 Cycloneda sanguinea
     Cycloneda sanguinea (Linnaeus) 

Genus Harmonia 

Multicolored Asian Lady Beetle - Harmonia axyridis   Red on Black 
lady beetle - Harmonia axyridis  Multicolored Asian Lady Beetle - Harmonia axyridis Lady beetle larva - 
Harmonia axyridis
     Harmonia axyridis (Pallas) - Non-native

Genus Hippodamia 

Convergent Ladybird Beetle for California in April - 
Hippodamia convergens convergent
 lady beetle? - Hippodamia convergens Convergent 
Lady Beetle - Hippodamia convergens Lady Beetle 
Larva ? - Hippodamia convergens Ladybird 
bettle swarm - Hippodamia convergens
     Hippodamia convergens Guérin-Méneville

Coccinellidae - Ladybird beetle - ?? - Hippodamia sinuata Coccinellidae - Ladybird beetle - ?? - Hippodamia sinuata
    Hippodamia sinuata crotchi Casey

Genus Mulsantina 

 Lady Beetle - Mulsantina picta Beetle - Mulsantina picta Painted Lady Beetle - Mulsantina picta Ladybug - Mulsantina picta Mulsantina picta? - Mulsantina picta
     Mulsantina picta (Randall)

Genus Myzia

Myzia interrupta (Casey) - Myzia interrupta Myzia interrupta (Casey) - Myzia interrupta Myzia interrupta (Casey) - Myzia interrupta
     Myzia interrupta (Casey) 

Lady Beetle - Myzia pullata Ladybird beetle - Myzia pullata Myzia pullata Larva of Unknown Lady Bug Species - Myzia pullata
     Myzia pullata (Say)

Genus Naemia 

Naemia seriata (Melsheimer) - Naemia seriata Ladybeetle - Naemia seriata Lady Beetle - Naemia seriata coccinellid larva - Naemia seriata
     Naemia seriata (Melsheimer) 

Genus Neoharmonia

Neoharmonia venusta venusta (Melsheimer) - Neoharmonia venusta Neoharmonia venusta venusta (Melsheimer) - Neoharmonia venusta Neoharmonia venusta venusta (Melsheimer) - Neoharmonia venusta Neoharmonia venusta venusta (Melsheimer) - Neoharmonia venusta Neoharmonia venusta What kind of ladybug is this? - Neoharmonia venusta
Neoharmonia venusta ampla (Mulsant) - Neoharmonia venusta Neoharmonia venusta ampla (Mulsant) - Neoharmonia venusta Neoharmonia venusta ampla (Mulsant) - Neoharmonia venusta Neoharmonia venusta ampla (Mulsant) - Neoharmonia venusta Neoharmonia venusta ampla (Mulsant) - Neoharmonia venusta Neoharmonia venusta ampla (Mulsant) - Neoharmonia venusta Ladybug - Neoharmonia venusta
    Neoharmonia venusta (Melsheimer)

Genus Olla 

Olla v-nigrum (Mulsant) - Olla v-nigrum Olla v-nigrum (Mulsant) - Olla v-nigrum Ashy Gray Ladybird Beetle - Olla v-nigrum Ashy Gray Ladybird Beetle - Olla v-nigrum Olla v-nigrum (Mulsant) - Olla v-nigrum Olla v-nigrum (Mulsant) - Olla v-nigrum  
     Olla v-nigrum (Mulsant) 

Genus Psyllobora

Psyllobora plagiata Schaeffer - Psyllobora plagiata
    Psyllobora plagiate Schaeffer

Psyllobora renifer Casey - Psyllobora renifer Psyllobora renifer Casey - Psyllobora renifer Psyllobora renifer Casey - Psyllobora renifer Psyllobora renifer Casey - Psyllobora renifer Psyllobora renifer Casey - Psyllobora renifer 
     Psyllobora renifer
Casey 

Psyllobora vigintimaculata (Say) - Psyllobora vigintimaculata Lady Beetle Larva ? - Psyllobora vigintimaculata
     Psyllobora vigintimaculata
(Say) 


Subfamily Epilachninae

Genus Epilachna 

Epilachna borealis (Fabricius) - Epilachna borealis ladybug - Epilachna borealis Very small spiked insect.  - Epilachna borealis
     Epilachna borealis (Fabricius) 

Epilachna tredecimnotata (Latreille) - Epilachna tredecimnotata Epilachna tredecimnotata (Latreille) - Epilachna tredecimnotata Epilachna tredecimnotata (Latreille) - Epilachna tredecimnotata
     Epilachna tredecimnotata (Latreille) 

Mexican bean beetle - Epilachna varivestis Mexican Bean Beetle  - Epilachna varivestis
     Epilachna varivestis Mulsant 


Selected Bibliography

Acorn, J. 2007. Ladybugs of Alberta, Finding the Spots and Connecting the Dots. University of Alberta Press, Edmonton. xxx + 169 pp.

Blaisdell, F.E. 1892. A new species of Coleoptera from California. Entomol. News 3: 51.

Blatchley, W.S. 1917. On some new or noteworthy Coleoptera from the west coast of Florida. Canadian Entomol. 49: 137-143.

Casey, T.L. 1899. A revision of the American Coccinellidae. Jour. New York Entomol. Soc. 7: 71-169.

Casey, T.L. 1908. Notes on the Coccinellidae, Canadian Entomol. 40: 393-421.

Chapin, J.B. 1973. New species of ladybeetles in the genera Scymnus and Mulsantina (Coleoptera: Coccinellidae). Ann. Entomol. Soc. America 66: 1071-1073.

Crotch, G.R. 1873. Revision of the Coccinellidae of the United States. Trans. American Entomol. Soc. 4: 363-382.

Crotch, G.R. 1874. Descriptions of new species of Coleoptera from the Pacific Coast of the United States. Trans. American Entomol. Soc. 5: 73-80.

DeGeer, C. 1775. Memoires pour Servir a l'Histoire des Insectes. Vol. 5. Stockholm, 448 pp.

Dobzhansky, T. 1941. Beetles of the genus Hyperaspis inhabiting the United States. Smithsonian Miscellaneous Collections, 101(6): 1-94.

Fabricius, J. C. 1775. Systema Entomologiae. Lipsiae, 832 pp.

Gordon, R.D. 1976. The Scymnini (Coleoptera: Coccinellidae) of the United States and Canada: Key to Genera and Revision of Scymnus, Nephus, and Diomus. Bulletin of the Buffalo Society of Natural Sciences 28: 1-362.

Gordon, R.D. 1985. The Coccinellidae (Coleoptera) of America north of Mexico. Journal of the New York Entomological Society 93(1): 1-912.

Gordon, R.D. and E.A. Chapin. 1983. A revision of the New World species of Stethorus Weise (Coleoptera: Coccinellidae). Trans. American Entomol. Soc. 109: 229-276.

Gorham, H.S. 1897. Biologia Centrali-Americana, Insecta, Coleoptera: Coccinellidae 7: 217-240.

Guerin-Meneville, F. E. 1829-1844. Iconographie du Regne Animal de Cuvier. Vol. 7, Insectes, 1829-1838. Paris, 576 pp.

Horn, G.H. 1895. Studies in Coccinellidae. Transactions of the American Entomol. Soc. 22: 81-114.

Latreille, P.A. 1833. Insectes de l'Amerique equinoxiale. Pp. 65-96 In: A. Humboldt and A. Bonpland, Recueil d' Observations de Zool ogle et d'Anatomie Comparee ; Faites dans l'Ocean Atlantique, dans l'Interieur du Nouveau Continent et dans la Mer du Sud, Pendant les Annees, 1799, 1800, 1801, 1802 et 1803, Vol. 2. Paris.

LeConte, J.L. 1850. General remarks upon the Coleoptera of Lake Superior. Pp. 201-242 In: Agassiz, Lake Superior: Its Physical Character, Vegetation and Animals, Compared with Those of Other and Similar Regions. Boston.

LeConte, J.L. 1852. Remarks upon the Coccinellidae of the United States. Proc. Acad. Nat. Sci. Philadelphia 6: 129-145.

LeConte, J.L. 1858. Descriptions of new species of Coleoptera, chiefly collected by the United States and Mexican Boundary Commission, under Maj. W. H. Emory, U.S.A. Proc. Acad. Nat. Sci. Philadelphia 10: 59-89.

LeConte, J.L. 1859. The Coleoptera of Kansas and eastern New Mexico. Smithsonian Cont. Knowl. 11: 1-58.

Linnaeus, C. 1758. Systema Naturae, 10th edition. Stockholm, 826 pp.

Linnaeus, C. 1763. Centuria Insectorum Rariorum. Uppsala, 32 pp.

Linnaeus, C. 1767. Systema Naturae, 12th edition. Stockholm, 830 pp.

Mannerheim, C.C. 1843. Beitrag zur Kafer-Fauna der Aleutischen Inseln, der Insel Sitka und Neu California. Bull. Soc. Imp. Nat. Moscou. 16: 175-314.

Melsheimer, F.E. 1847. Descriptions of new species of Coleoptera of the United States. Proc. Acad. Nat. Sci. Philadelphia 3: 158-181.

Mulsant, M.E. 1850. Species de Coleopteres trimeres securipalpes. Ann. Sci. Phys. Nat. Lyon 2: 1-1104.

Olivier, A.G. 1808. Entomologie, ou Histoire Naturelle des Insectes, avec leurs Caractcres Generiques et Specifiques, leur Description, leurs Synonymie, et leurs Figures Enluminee. Coleopteres, Vol. 5. Paris.

Randall, J.W. 1838. Description of New Species of Coleopterous Insects Inhabiting the State of Maine. Boston Society of Natural History, 2(1): 1-33.

Say, T. 1824. Narrative of an expedition to the source of the St. Peter's River, etc., under the command of Stephen H. Long, Major, U.S.T.E. Jour. Acad. Nat. Sci. Philadelphia 2: 268-378.

Say, T. 1826. Descriptions of new species of coleopterous insects inhabiting the United States. Jour. Acad. Nat. Sci. Philadelphia 5: 238-284, 293-304.

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